First of - WHAT THE HELL ARE SHUGGURATHS?!? They are amazingly cool and dare I say beautiful!
Definitely very Lovecraftian - Cthulhu inspired!
But why are they on Elpis?
And not Pandora?
Do the Ratthyds come from inside them?
At least with Kraggons you know they are born of rock and fire.
Next, are Threshers possibly related to the Destroyer somehow (tentacles)?
Could they have come into being on Elpis somehow when Jack brought the Eye and who knows what other parts of the Destroyer up to Elpis?
Nakayama refers to Meg the Thresher, as well as the Tiny Destroyer, as genetic abominations.
Maybe threshers are a Hyperion R&D experiment that got loose and found their way down to Elpis?
Or were deliberately put there by Hyperion?
Also, I love how Swagmen have those unique, extra tough flies with them.
Finally, Torks should evolve like Varkids, damn it!